Joël Lodé
Taxonomy of the Cactaceae. Volumes 1-2. 2015.
Lodé's classification of cacti is based, among other things, on new molecular data and their interpretation. The Genera Acharagma, Escobaria, Neobesseya, Ortegocactus and Pelecyphora are listed separately in this publication.
Genus Acharagma
| Diagnosis of Genus: see description of genus | |
| Currently 3 recognised species, and one doubtful subspecies: | |
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Species:
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| Acharagma aguirreanum (Glass et Foster) Glass, 1998 | |
| Synonyme: | Gymnocactus aguirreanus |
| Escobaria aguirreana | |
| Thelocactus aguirreanus | |
| Acharagma huasteca Elhart, 2011 | |
| Acharagma roseanum (Boedeker) Anderson, 1999 | |
| Synonyme: | Gymnocactus roseanus |
| Coryphantha roseana | |
| Escobaria roseana | |
| Neolloydia roseana | |
| Thelocactus roseanus | |
| Acharagma roseanum subsp. galeanense (Haugg) Hunt, 2002 | |
| Synonyme: | Escobaria roseana subsp. galeanensis |
| Notes: | A. roseanum subsp. galeanense does not seem to be in its right place, and could be properly considered as a separate species. In my option (Lodé), it s a taxon closer to the recently discovered A. huasteca. |
Genus Escobaria
| Diagnosis of Genus: see description of genus | |
| Currently 9 recognised species (+3 subspecies): | |
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Species:
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| Acharagma albicolumnaria Hester, 1941 | |
| Synonyme: | Coryphantha albicolumnaria |
| Acharagma alversonii (J. M. Coult.) N. P. Taylor, 1997 | |
| Synonyme: | Escobaria vivipara var. alversonii |
| Coryphantha alversonii | |
| Coryphantha vivipara var. alversonii | |
| Escobaria chihuahuensis Briton et Rose, 1923 | |
| Synonyme: | Coryphantha chihuahuensis (?) |
| Escocoryphantha chihuahuensis | |
| Notes: | possibly wrongly placed |
| Escobaria chihuahuensis subsp. henricksonii (Glass et Foster) N. P. Taylor, 1998 | |
| Synonyme: | Coryphantha henricksonii (?) |
| Escobaria henricksonii (?) | |
| Escocoryphantha henricksonii | |
| Notes: | possibly wrongly placed |
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Escobaria desertii (Engelm.) Buxb., 1951 [ex Coryphantha chlorantha (Engelm.) Br. et R.] |
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| Synonyme: | Coryphantha chlorantha |
| Coryphantha chlorantha var. deserti | |
| Escobaria chlorantha | |
| Escobaria hesteri (Y, Wright) Buxb., 1951 | |
| Synonyme: | Coryphantha hesteri |
| Escobaria laredoi (Glass et Foster) N. P. Taylor, 1979 | |
| Synonyme: | Coryphantha laredoi |
| Escobaria rigida | |
| Notes: | possibly wrongly placed |
| Escobaria sneedii Britton et Rose, 1923 | |
| Synonyme: | Coryphantha sneedii |
| Coryphantha pygmaea | |
| Notes: |
The following taxa were included among the synonyms of E. sneedii in the Taxonomy of the Cataceae, Alphabetical index of taxa in current usage and their synonyms (corrigenda & addenda at the end of the index) / version online 3.9 (2015): -Coryphantha orcuttii -Coryphantha organensis -Coryphantha pygmaea -Coryphantha strobiliformis var. orcuttii -Escobaria guadalupensis -Escobaria leei -Escobaria orcuttii -Escobaria orcuttii var. koenigii -Escobaria orcuttii var. macraxina -Escobaria organensis -Escobaria sandbergii -Escobaria villardii |
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Escobaria sneedii subsp. leei (Boed.) D. R. Hunt, 1997 |
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| Synonyme: | Coryphantha sneedii var. leei |
| Escobaria leei | |
| Escobaria sneedii subsp. orcuttii (Boed.) Luethy, 1999 | |
| Synonyme: | Coryphantha orcuttii |
| Escobaria orcuttii var. koenigii | |
| Escobaria orcuttii var. macraxina | |
| Neolloydia orcuttii | |
| Escobaria tuberculosa (Engelm.) Britton et Rose, 1923 | |
| Synonyme: | Coryphantha tuberculosa |
| Coryphantha tuberculosa var. durispina | |
| Coryphantha strobiliformis | |
| Coryphantha strobiliformis var. durispina | |
| Escobaria strobiliformis | |
| Coryphantha varicolor | |
| Escobaria varicolor | |
| Escobaria vivipara (Nutt.) Buxb., 1951 | |
| Synonyme: | Coryphantha alversonii var. exaltissima |
| Coryphantha arizonica | |
| Coryphantha columnaris | |
| Coryphantha neo-mexicana | |
| Coryphantha oklahomensis | |
| Coryphantha radiosa | |
| Coryphantha rosea | |
| Coryphantha vivipara | |
| Coryphantha vivipara var. aggregata | |
| Coryphantha vivipara var. alversonii (synonymum - probably a mistake, note - Jan Mynář) | |
| Coryphantha vivipara var. arizonica | |
| Coryphantha vivipara var. bisbeeana | |
| Coryphantha vivipara var. buoflama | |
| Coryphantha vivipara var. deserti (synonymum - probably a mistake, note - Jan Mynář) | |
| Coryphantha vivipara var. kaibabensis | |
| Coryphantha vivipara var. neo-mexicana | |
| Coryphantha vivipara var. radiosa | |
| Escobaria arizonica | |
| Escobaria bisbeeana | |
| Escobaria neomexicana | |
| Escobaria radiosa | |
| Escobaria bisbeeana | |
| Escobaria vivipara var. alversonii (synonymum - probably a mistake, note - Jan Mynář) | |
| Escobaria vivipara var. arizonica | |
| Escobaria vivipara var. bisbeeana | |
| Escobaria vivipara var. buoflama | |
| Escobaria vivipara var. deserti (synonymum - probably a mistake, note - Jan Mynář) | |
| Escobaria vivipara var. kaibabensis | |
| Escobaria vivipara var. neomexicana | |
| Escobaria vivipara var. radiosa | |
| Escobaria vivipara var. rosea | |
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Taxonomy of the Cataceae (supplement) Alphabetical index of taxa in current usage and their synonyms (corrigenda & addenda at the end of the index) / version online 3.9 (2015) Recognised taxa and omitted in the Volume I: |
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Escobaria hesteri subsp. grata |
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Genus Neobesseya
| Diagnosis of Genus: see description of genus | |
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Die Gattung Neobesseya wird in dieser Arbeit als korrekt angesehen, einschließlich Taxa, die in der Sektion Neobesseya von Escobaria nach Taylor (in The New Cactus Lexicon, 2006). Diese Einstufung (basierend auf der Arbeit von B. Magnani proparte 2011) wird nicht durch eine molekularbiologische Analyse der betreffenden Art gestützt und ist daher vorbehaltlos zu nehmen.
Currently 10 recognised species + 2 subspecies: |
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Species:
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Neobesseya abdita (Řepka et Vaško) J. Lodé, 2013 (ex Escob. abdita Řepka et Vaško 2011) |
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| Synonyme: | Escobaria abdita |
| Neobesseya cubensis (Britton et Rose) Hester, 1941 | |
| Synonyme: | Coryphantha cubensis |
| Escobaria cubensis | |
| Neolloydia cubensis | |
| Neobesseya dasyacantha (Engelm.) J. Lodé, 2013 | |
| Synonyme: | Coryphantha dasyacantha |
| Coryphantha dasyacantha var. varicolor (as synonym for Neobesseya) | |
| Escobaria dasyacantha | |
| Escobaria dasyacantha var. varicolor | |
| Escobaria duncanii (note - Jan Mynář: probably a mistake, is synonym of Neob. duncanii) | |
| Escobesseya duncanii (note - Jan Mynář: probably a mistake, is synonym of Neob. duncanii) | |
| Escobesseya dasyacantha | |
| Neobesseya dasyacantha subsp. chaffeyi (Britton et Rose) J. Lodé, 2013 | |
| Synonyme: | Escobaria chaffey (as synonym of E. dasyacantha) |
| Escobaria dasyacantha var. chaffey | |
| Neobesseya duncanii (Hester) J. Lodé, 2013 | |
| Synonyme: | Coryphantha dasyacantha var. duncanii |
| Coryphantha duncanii | |
| Escobaria duncanii (as synonym of E. dasyacantha; probably a mistake - note of Jan Mynar) | |
| Escobesseya duncanii (as synonym of E. dasyacantha; probably a mistake - note of Jan Mynar) | |
| Neobesseya emskoetteriana (Quehl) J. Lodé, 2013 | |
| Synonyme: | Coryphantha emskoetteriana |
| Coryphantha muehlbaueriana | |
| Coryphantha piercei | |
| Coryphantha roberti | |
| Escobaria bella | |
| Escobaria emskoetteriana | |
| Escobaria muehlbaueriana | |
| Escobaria runyonii | |
| Neobesseya muehlbaueriana | |
| Neobesseya lloydii (Britton et Rose) J. Lodé, 2013 | |
| Synonyme: | Escobaria lloydii |
| Neobesseya minima (Baird) J. Lodé, 2013 | |
| Synonyme: | Coryphantha minima |
| Coryphantha nellieae | |
| Escobaria minima | |
| Escobaria nellieae | |
| Neobesseya missouriensis (Sweet) Britton et Rose, 1923 | |
| Synonyme: | Coryphantha marstonii |
| Coryphantha marstonii | |
| Coryphantha missouriensis var.marstonii | |
| Coryphantha similis | |
| Escobaria missouriensis | |
| Escobaria missouriensis subsp. navajoensis | |
| Escobaria missouriensis var. caespitosa | |
| Escobaria missouriensis var. marstonii | |
| Escobaria missouriensis var. similis | |
| Neobesseya notesteinii | |
| Neobesseya roseiflora | |
| Neobesseya similis | |
| Neobesseya wissmannii | |
| Neobesseya missouriensis subsp. asperispina (Boedeker) J. Lodé, 2013 | |
| Synonyme: | Coryphantha asperispina (as synonym for Neobesseya missouriensis) |
| Escobaria asperispina (as synonym for Neobesseya missouriensis) | |
| Escobaria missouriensis subsp. asperispina | |
| Neobesseya asperispina | |
| Neobesseya robbinsorum (W. Earle) Doweld, 2000 | |
| Synonyme: | Cochiseia robbinsorum |
| Coryphantha robbinsorum | |
| Escobaria robbinsorum | |
| Neobesseya zilziana (Boed.) Boed., 1933 | |
| Synonyme: | Coryphantha zilziana |
| Escobaria zilziana | |
Genus Ortegocactus
| Diagnosis of Genus: see description of genus | |
| Currently only 1 recognised species: | |
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Species:
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| Ortegocactus macdougallii Alexander, 1961 | |
| Synonyme: | Neobesseya macdougallii |
Genus Pelecyphora
| Diagnosis of Genus: see description of genus | |
| Currently 2 recognised species: | |
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Species:
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| Pelecyphora aselliformis Ehrenb., 1843 | |
| Synonyme: | Pelecyphora aselliformis var. concolor |
| Pelecyphora aselliformis var. grandiflora | |
| Ariocarpus aselliformis | |
| Pelecyphora strobiliformis (Werderm.) Frič et Schelle, 1935 | |
| Synonyme: | Ariocarpus strobiliformis |